Table 1 Species of haemogregarines of the genus Hepatozoon described from African varanids

Hepatozoon borreli (Nicolle & Comte, 1906) Smith, 1996^b

Varanus griseus (Daudin, 1803)

Tunisia

7–8 × 2/1–3 × 1–2

[3]^b, [6, 14]

Hepatozoon camarai (Dias, 1954) Smith, 1996^b

Varanus albigularis albigularis (Daudin, 1802)

Mozambique

2 forms observed: banana-shaped: 11.75 × 5.00; long curved: 14.25–18.25 × 1.25–4.25/nucleus irregular

[3]^b, [9]

Hepatozoon paradoxa (Dias, 1954) Smith, 1996^b

V. a. albigularis ¹

Mozambique

V. niloticus (Kenya)²; V. a. albigularis and V. niloticus (South Africa)³

6.75–7.50 × 4.25–5.50/nucleus not visible¹; 8.1 × 5.2/nucleus irregular or not visible²; 6.99 × 4.39/nucleus not visible³

[9]¹, [30]², this study³, [3]^b

Haemogregarina roshdyi Ramdan, Sauod, Mohammed & Fawzi, 1996 [probably Hepatozoon roshdyi (Ramdan, Sauod, Mohammed & Fawzi, 1996)]

V. griseus

Egypt

13–20 × 1.5–2.5/6.0–8.5 × 1.5–2.5

[40]

^a Hepatozoon toddi (Wolbach, 1914) Smith, 1996^b

Varanus niloticus (Linnaeus, 1762)

The Gambia

10.3 × 2.5/not given

[3]^b, [6, 15]

Hepatozoon varani (Laveran, 1905) Smith, 1996^b

V. niloticus ¹

South Africa

V. niloticus and V. griseus (Senegal)²; V. niloticus (?) (Portuguese Guinea)³; V. niloticus (The Gambia)⁴; V. niloticus (Senegal)⁵; Varanus arenarius Duméril & Bibron, 1836 (?) (French West Africa)⁶; V. griseus (French Sudan)⁷; V. niloticus (Kenya)⁸

14 × 3/not given¹; 11–15 × 3/not given²; 12.0 × 13/3.5–4.5 × 3.0³; 10.3 × 2.5/not given^a4; two forms: 12–14 × 2/not given, 10–12 × 4–5/5–6 × 4–5⁵; report only⁶; report only⁷; 12.7 × 4.6/not given⁸

[13]¹, [41]², [15]³, [42]³, [43]⁴, [44]⁵, [45]⁶, [46]⁷, [30]⁸, [3]^b