Fig. 1

Study area. The two studied communities within Loja Province in southern Ecuador are depicted. They are separated by approximately 28.3 km. Circle diameter is proportional to sample size in each locality. Map generated with ArcGIS software, version 10.5, based on data freely available from the Military Geographic Institute of Ecuador (IGM, http://www.geoportaligm.gob.ec/portal/). Community location was obtained via GPS measurement